It’s Claire’s 60th Birthday and Betty has to deal with Ignacio and Elena possibly dating. Meanwhile, Wilhelmina and Daniel are jealous of Conner and Molly’s memorable relationship.

Betty comes up with a solution to deal with her personal problems, but her crush on her cute neighbor Jesse is keeping her distracted. Meanwhile, Connor tries to track down who was on vacation with Molly and Daniel desperately tries to cover his tracks.

Betty and Hilda fight over how they should take care of Ignacio. Daniel tries to cover up evidence of his vacation with Molly. Connor and Wilhelmina can’t keep their hands off each other at the office.

Nikki Blonsky guest stars as Terri, Betty’s frenemy that works at rival magazine Elle.

Betty and Amanda are forced to work together for a Mode website article “How I Blew Ten Grand Without Spending a Dime.” The two girls go out for a night on the town for research and Betty is forced to do something very Amandaish.

Betty and Marc compete in the same program, The Young Editors Training (YETI). Wilhelmina, Connor, Marc and Molly have a weird love rectangle going on. Justin and Randy grow close.

Betty needs Daniel to help prevent a PR nightmare at Mode, but he’s stuck at a company retreat. Meanwhile, Hilda runs an illegal beauty salon when the city denies her permit for a legal business and Connor introduces his fiance to Mode.

Betty invites Jesse’s band to play at the Mode after party, Cliff asks Marc to move in, Amanda crashes at Betty’s, and Wilhelmina and Daniel fight for Connor to join their side.

Betty finds herself in an all out war with Kimmie Keegan (Lindsay Lohan) and Ignacio gets ready to vote for the very first time.  Hilda takes advantage of election traffic to promote her business and Wilhelmina tries to hire a financial officer Daniel hates.

In Granny Pants, Betty’s old enemy, Kimmie Keegan is back! Betty gets Kimmie a job at Mode and the two look like they’re going to be friends.