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Desperate to help his pregnant daughter, Mark follows Addison to Oceanside. Derek's suspicions are raised when the Chief recruits Meredith to assist with a high profile operation.
Thanksgiving, Christmas and New Year's pass. Miranda is visited by her father. Mark and Lexie cope with the arrival of a woman from his past. Thatcher questions the Chief's behavior.
Izzie returns to Seattle Grace with an important figure from her past. Stress takes a toll on Richard. Owen recruits a fellow Iraq war vet as the a cardiothoracic surgeon.
All eyes are on Arizona to help a patient with an enormous donation to spare. Cristina tries to test Owen's boundaries, and Alex is losing it over Izzie.
Derek must help the most unlikeliest of all; the hospital lab-tech, Isaac, when it turns out that he has an inoperable tumor surrounding his spine. McDreamy also questions Richard's authority.
A burn victim's death in a chaotic ER prompts the Chief and the board to find who is responsible, as all the Mercy West and Seattle Grace residents scramble to save their jobs.
Newly-merged Seattle Grace and Mercy West residents vie for surgeries and their careers, while Arizona stands by Callie when her father employs a priest to convince her to date men, and Izzie experiences the brunt of her new competition.
When Meredith and Lexie Grey's father, Thatcher, returns to the hospital with a failed liver, it's up to Meredith to save his life. That won't be easy for myriad reasons.
Izzie returns to work after a near-death experience. An announcement from the Chief has residents expecting layoffs and scrambling to survive the cuts.
The doctors of Seattle Grace continue to battle their own demons and cope with the loss of George in their own way, while a couple of patients make a big impact on the doctors, too.