Wilhelmina Slater is a diva and former supermodel who works as the Creative Director of Mode.  However, she will do whatever it takes to get the role of Editor in Chief that she feels was unfairly given to Daniel.  She does claim it for a bit in season 3 but loses the title again by the end of episode 2.  She is now co-Editor in Chief alongside Daniel.

Wilhelmina Slater Quotes

Who cares what the average woman wears on the street. We shouldn't be taking pictures of them, we should be throwing rocks at them.

Wilhelmnia

The rounded o's of a stunted adolescent. The stiff t's of someone who is clearing overcompensating. This is Daniel's handwriting.

Wilhelmnia
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Ugly Betty Quotes

You know exactly what [Whilemnia] is up to. Firing, scheming, looking for a puppy to kick.

Betty

Betty: Does every spread have to be women in skimpy bras?
Daniel: You're right, I haven't thought of that. Let's lose those bras please.